Inés,
> in order to decide whether something succeeded or not in producing copies, I need a notion of equality, or similarity.. don't you?
Well, I recall there are complex error-correcting mechanisms that check on RNA and repair mistakes, as well as splicing out introns, etc. But at a basic level, in biology, I think that success means ongoing reproductive success, and accurate copying only serves that end. After all, in sexually reproducing species, offspring are not genetic copies of the parents. Most of the genes need to be exact copies for the offspring to survive and reproduce, but ultimately it's their reproductive success that "decides" whether any member of a species is a good enough copy... so far as future evolution is concerned.
> I always fall back on the need to try to define "copy" and "mistake", otherwise the recursive definition seems to dilute away in "something affecting something else".
In biology, the key thing that has to get reproduced is the ability to reproduce. With us humans, the key thing that needs to be communicated is the ability to communicate. The latter is much more complicated, because only in special cases (like learning a math formula or relaying news) does it come down to accurate copying. Even then, the meaning of the news or the formula depends on what we do with it in future.
Your bus example is good... it shows how meaning ramifies out into the world to have all kinds of effects. Likewise in biology, the successful reproduction of an organism can cause many different effects in the world. But the key effect is to make more reproduction possible. With communication, the key function is to keep the conversation going, whether with yourself or with another person... to keep making new things seem fun or important to talk and think about. You're right that "sameness" is a crucial element, though less clearly defined than in biology. If you and I had very different notions, we would just be talking past each other. But again, while accurately grasping each other's thoughts is important, it serves a deeper purpose - for me anyway, it makes me feel much more connected and hopeful, maybe better able to listen and express myself clearly in future.
In short, "meaning" does indeed dilute out into the world of interaction... I think that's why it can seem impossible to define except as a feeling we have about things. Meaning does more than set up the possibility for more meaning, but that's the key thing it succeeds or fails at.
Hope that clarifies things a little... and thanks again for your helpful responses.
Conrad